整个完整的句子应该是This program will tell you how what you have learned can be used in …… 吧
这里的意思是：这个项目会告诉你（使你明白）,你所学到的东西能够怎么样的被运用到……里面
首先,tell you sth.后面整个句子充当sth.
tell you how sth can be used in …….告诉你sth.如何能被运用到……里面去
而这里是用整个句子what you have learned 充当 sth.
what you have learned what 在句子中充当宾语