求牛人帮忙解析下如下句子的含义Which of the following,if true.best explains how i_英语解答

求牛人帮忙解析下如下句子的含义
Which of the following,if true.best explains how improvements in pesticide use have been accompanied by greater losses to certain pests?
请详细解析下how在这的用法,

人气：363 ℃　时间：2020-03-25 07:05:41

解答

how 在这里引导宾语从句,作explains 的宾语.
句子的主干结构:
Which (主语) ...explains (谓语) how (整个从句作宾语)...
在宾语从句中,improvements 主语 have been accompanied 谓语 how 作状语.