求牛人帮忙解析下如下句子的含义
Which of the following,if true.best explains how improvements in pesticide use have been accompanied by greater losses to certain pests?
请详细解析下how在这的用法,
人气：338 ℃ 时间：2020-03-25 07:05:41
解答
how 在这里引导宾语从句,作explains 的宾语.
句子的主干结构:
Which (主语) ...explains (谓语) how (整个从句作宾语)...
在宾语从句中,improvements 主语 have been accompanied 谓语 how 作状语.
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